| name | Amanita valens | ||||||||||||||||||||
| author | (E.-J. Gilbert) Bertault. 1980. Bull. Trimestriel. Soc. Mycol. France 96(3): 281. [Misapplication to Amanita curtipes E.-J. Gilbert.] | ||||||||||||||||||||
| name status | nomen acceptum | ||||||||||||||||||||
| synonyms |
≡Amidella lepiotoides f. valens E.-J. Gilbert nom. nud. 1940. Iconogr. Mycol. (Milan) 27, suppl. 1 (1): 77, 126, tab. 23 (fig. 4-5). [Lacking Latin diagnosis, specification of holotype, etc. ICBN §36.1, §37.1, etc.]
≡Amanita lepiotoides f. valens E.-J. Gilbert. 1941. Iconogr. Mycol. (Milan) 27, suppl. (2): 291, tab. 31. [Parrot (1960. Amanites S.-O. France: 58) appears to mistakenly propose this combination as new—"fo. valens Gilbert, nova forma."]
≡Amanita valens (E.-J. Gilbert) Kühner & Romagn. nom. inval. 1954. Fl. Anal. Champ. Supér.: 431. [Lacking full and direct reference to basionym. ICBN §33.2]
≡Amanita curtipes var. valens (E.-J. Gilbert) Contu. 2000. Boll. Gruppo Micol. G. Bresadola 43(2): 84.
≡Amanita ponderosa f. valens (E.-J. Gilbert) Neville & Poumarat. 2004. Fungi Europaei 9: 691. The editors of this site owe a great debt to Dr. Cornelis Bas whose famous cigar box files of Amanita nomenclatural information gathered over three or more decades were made available to RET for computerization and make up the lion's share of the nomenclatural information presented on this site. | ||||||||||||||||||||
| MycoBank nos. | 112740, 345951, 351646, 467412, 366240 | ||||||||||||||||||||
| GenBank nos. |
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| lectotypes | E.-J. Gilbert. 1941. Iconogr. Mycol. (Milan) 27 suppl. 1 (3): pl. 31. | ||||||||||||||||||||
| neotypes | A neotype was incorrectly proposed (original material is extant ICBN §9.6) for this taxon by Neville & Poumarat (2004). The proposed specimen is in herb. S. Poumarat. | ||||||||||||||||||||
| lectotypifications | Tulloss. 2005a. Mycotaxon 92: 481. | ||||||||||||||||||||
| intro |
Olive text indicates a specimen that has not been thoroughly examined (for example, for microscopic details) and marks other places in the text where data is missing or uncertain. The following material not directly from the protolog of the present taxon is from original research of Dr. C. Bas (collections and notes in L) and R. E. Tulloss. | ||||||||||||||||||||
| pileus context | filamentous undifferentiated hyphae 1.5 - 7.5 µm wide, branching, sometimes in fascicles; acrophysalides globose to subglobose to broadly ellipsoid to broadly clavate to clavate to pyriform, up to 83 × 61 µm, thin-walled, dominant; vascular hyphae 2.0 - 3.5 µm wide. | ||||||||||||||||||||
| subhymenium | cellular in mature material comprising rather small globose to subglobose to broadly ellipsoid inflated cells (up to 16 × 13 µm) in chains forming 2 - 3 layers, subcellular in immature material comprising clavate to ventricose subinflated cells and uninflated short hyphal segments (indicating subhymenial cells probably inflate late in expansion of basidiome as in many amanitas). | ||||||||||||||||||||
| basidia | 40 - 56 × 9.6 - 11.8 µm, dominantly 4-, but occasionally 2-spored, thin-walled, clavate; no clamps observed. | ||||||||||||||||||||
| universal veil | On pileus: absent. On stipe base, exterior surface layer: about 45 μm thick, thinner in some areas; filamentous undifferentiated hyphae 4.4 - 5.4 µm wide, interwoven gelatinizing; vascular hyphae present, 2.0 - 2.5 µm wide. On stipe base, subsurface layer: inflated cells thin-walled, up to 28 × 12 µm, ellipsoid to elongate accompanied by slightly inflated short hyphal segments. On stipe base, interior: filamentous undifferentiated hyphae 3.2 - 9.3 µm wide, interwoven; inflated cells plentiful, elongate to clavate (up to 24 × 15 µm, with walls thin to 0.7 µm thick) or subglobose to ellipsoid (up to 104 × 81 µm, with walls thin to 1.1 µm thick); vascular hyphae present, 1.4± µm wide. On stipe base, inner surface: 3 - 7 hyphal diameters thick; filamentous undifferentiated hyphae partially gelatinizing, 1.7 - 4.4 µm wide, longtiudinally oriented, with some swollen terminal segments short subclavate; vascular hyphae not observed. | ||||||||||||||||||||
| stipe context | longitudinally acrophysalidic; filamentous undifferentiated hyphae 1.1 - 5.6 µm wide, branching, sometimes with slightly thickened, slightly yellowish walls; acrophysalides dominating, up to 192 × 37 µm, thin-walled or with walls to 1.0 µm thick and yellowish; vascular hyphae 3.2 - 4.2 µm wide, branching, uncommon. | ||||||||||||||||||||
| partial veil | absent. | ||||||||||||||||||||
| lamella edge tissue | not examined. | ||||||||||||||||||||
| basidiospores | composite of data from all material revised by RET: [140/6/4] (7.8-) 9.5 - 13.5 (-16.5) × (5.5-) 5.8 - 7.5 (-10.0) µm, (L = 10.5 - 12.5 µm; L’ = 11.6 µm; W = 6.3 - 7.2 µm; W’ = 6.6 µm; Q = (1.33-) 1.50 - 2.03 (-2.21); Q = 1.67 - 1.95; Q’ = 1.77), amyloid, thin-walled, smooth, hyaline, ellipsoid to elongate to cylindric, occasionally adaxially flattened, often expanded at one end; apiculus proportionately small, sublateral, cylindric; contents granular to guttulate; probably whitish in deposit. | ||||||||||||||||||||
| ecology | Subgregarious. Corsican collection in October under Pinus at 400± m elev. Marchand (1973) who seems to have collected material fitting the original description of Gilbert (1941) says the range of this species is from the Atlantic coasts of Spain and Portugal to the Mediterranean and south to Tanger. We must be cautious of the latter because of the uncertainity of Bertault’s concept of the species. Marchand further states that the species is found in spring and autumn and, further south, in winter, under Quercus and in mixed woods containing Pinus and deciduous trees; he cites one collection in September at Calmeilles, Pyrenées Orientales, France under Juniperus, Cistus, and scattered Q. suber (chêne liège). | ||||||||||||||||||||
| material examined | FRANCE: CORSICA—Unkn. Subdiv. - Forêt de Bonifatu, Calenzana, 29.x.1982 J. Häfner s.n. (L). SPAIN: ÁVILA— Unkn. Subdiv. - Ávila, 17.v.1974 G. Lopes s.n. (L). MADRID—Unkn. Subdiv. - Madrid, 6.v.1974 Sr. Luciano s.n. (L). TOLEDO—Unkn. Subdiv. - Naverhermosa, 5.v.1974 Srs.? de Amanategui s.n. (L). | ||||||||||||||||||||
| discussion |
Castro (1998. Mycotaxon 67: 231) proposed that the name of the present species was a synonym of Amanita curtipes E.-J. Gilbert. The following diagram compares sporographs of the two species: Notice that using spore size and shape to segregate the taxa seems to work rather well. | ||||||||||||||||||||
| citations | —R. E. Tulloss | ||||||||||||||||||||
| editors | RET | ||||||||||||||||||||
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| name | Amanita valens |
| bottom links | [ Keys & Checklists ] |
| name | Amanita valens |
| bottom links | [ Keys & Checklists ] |
Spore data sets and their composite
Each spore data set is intended to comprise a set of measurements from a single specimen made by a single observer; and explanations prepared for this site talk about specimen-observer pairs associated with each data set. Combining more data into a single data set is non-optimal because it obscures observer differences (which may be valuable for instructional purposes, for example) and may obscure instances in which a single collection inadvertently contains a mixture of taxa.
Text and User-Generated Sporographs are published under the Creative Commons License.
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